Here's where I recommend you start:
This is the best newbie-focused video series I have found, it was made before guides, were implimented, so the item tutorial isn't as important, but the information is still incredibly useful: http://www.youtube.com/playlist?list=PLDdv1UhKaheYX_f-7GYbx5uKjF8SheYnP
I recommend shying away from some of the more pro-oriented videos from people like Purge. They will throw around so many terms and acronyms that you will probably get overwhelmed and discouraged. DotaCinema has some good video guides, particularly the "hero in 60 seconds" series.
Complete the tutorial, all of it, including the 10 games with other players. For bonus points, do do the "last hit" tutorial. Learning how to last hit is what will allow you to play a carry instead of being in support hell.
Play "limited hero" matches until you have played every hero and can roughly identify what their role is in the game. Learning how each hero works is probably one of the more important differences when compared to LoL. In LoL, there are a few cookie cutter hero styles that most heroes are cut from. In DotA, there are some incredibly unique heroes that if you aren't aware of how they work, will kill you immediately (for example, if bloodseeker uses his ult on you, you will die if you run away.) The limited pool will keep you from feeding while you build your knowledge.
Join the GiantBomb chat channel in game. Specifically look for the term "LL MM". This means Low Level Matchmaking and will be newer players grouping up. You can also say in there that you are new and would like to have a mentor or someone help you learn. Bot matches are also good, but they can be frustrating. The bots tend to get into a 5-man group fairly early on, and without cooperation from your human teammates, they can easily overwhelm you.
Get a mic and use it, I can't emphasize how important this is. Even if you're new, as long as you are friendly and vocalize your concerns, this goes a long way to ensuring you will get help and not get yelled at. When someone asks you to place wards, ask "where is a good spot"? and they will ping on the map for you. If you are following a guide, it's ok to ask if the item is good or not. "This guide says to build a Mekanism, is that a good idea?" "No, I'm already building one and there's no reason to have two on a team, build a Force Staff instead". If someone says that as a response, friend them immediately. :p
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