You lose almost everything, including CoD points. Things like weapons, attachments, camouflage, pretty reticles, lethal, tactical and equipment items, facepaints, killstreaks, all perks, all pro versions of perks, challenges (not lifetime challenges), kill streaks all disappear
You retain:
Playercard pictures/layers/backgrounds.
Your emblem (the one that goes on your playercard)
Clan Tag
Lifetime challenges
Lifetime stats
So, spend your money on buying layers, or playercard related things before you prestige.
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