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Colonel Yevgeny Volgin is the leader of the GRU, and the secondary antagonist of Metal Gear Solid 3: Snake Eater.
Gene (formerly FOX codename Viper) is the main antagonist of Metal Gear Solid: Portable Ops. He is a key device to the series, offering Big Boss both the methods and means of constructing Outer Heaven.
Revolver Ocelot, aka Shalashaska, real name Adam, appears in the the Metal Gear Solid series with often unclear motives.
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